首先，他们考虑了有毒的或者无氧的底部水流的季节性上升造成这种灾难的可能性。

First, they considered the possibility that seasonal upwellings of toxic or oxygen-poor bottom water were responsible.

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从甲酸盐中获取所需的能量是目前最简单的维持生命方式，虽然它依赖于无氧的环境。

Getting useful energy from formate is now the simplest way known for any life form to get its fuel, though it relies on the absence of oxygen.

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因此，脊椎动物的无氧能量储备与动物的大小成正比。

Thus the anaerobic energy reserves of a vertebrate are proportional to the size of the animal.

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