If evolution is survival of the fittest, nature red in tooth and claw, how could animals evolve moral feelings?

如果进化论是讲适者生存，在弱肉强食的自然界，动物怎么能进化出道德情感？

耶鲁公开课 - 心理学导论课程节选

If both religion and reason are removed, all that remains is will and power, where the only law is the law of tooth and claw.

如果宗教和理性都被世人所摒弃,那么世间就只剩下了权力和欲望,而这世间唯一准则就只剩下了弱肉强食

耶鲁公开课 - 古希腊历史简介课程节选