对于结合，免疫球蛋白直接吸附在颗粒的表面上，主要通过London-范德华力和疏水相互作用。
For conjugation, immunoglobulin is directly adsorbed on the particle surfaces, mediated mainly by London–van der Waals force and hydrophobic interaction.
四聚物的结合是通过表面免疫球蛋白抗原受体，通过Fc受体结合抗体的很少量甚至没有。
Binding of the tetramers occurred by the surface immunoglobulin antigen receptor with little or no binding to antibody captured in the Fc receptor.
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