珠被内层细胞特化为珠被绒毡层。
Inner cells in the integument differentiated into integument tapeta.
大胞子四分体线形排列，合点端大孢子为功能大孢子，胚囊发育类型为蓼型，具有特化的珠被绒毡层和承珠盘。
Megaspore tetrad is linear. the chalazal megaspore develops into an embryo sac of polygonum type. Hypostase and special integument tapetum exist in the ovule.
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