The key question of what caused the leak remains unanswered.

泄漏究竟是怎么造成的，这一关键问题仍未得到答案。

《牛津词典》

What caused the monsoon to migrate?

是什么导致了季风的迁移呢？

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Future research should look at the behavior and ecology of these worms, to understand what caused the new species to arise.

未来的研究应该看看这些蠕虫的行为与生态学，以了解新物种的出现是由什么造成的。

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But he said it was too early to say what caused the April twentieth explosion on the Deepwater Horizon drilling rig.

VOA: special.2010.06.19

The semantics was what caused the problem, because the operator was expecting a particular kind of structure there.

引起问题的是语义，因为运算符需要的是,一种特定种类的数据结构。

麻省理工公开课 - 计算机科学及编程导论课程节选

Milton, of course, is instructing the muse to explain first, before she gets around to explaining anything else, what caused Adam and Eve to fall.

弥尔顿当然是在让缪斯先说，在她说其他事情之前，是什么让亚当夏娃堕落的。

耶鲁公开课 - 弥尔顿课程节选

Predictably, this was what caused the harsh words in the Delhi talks this week.

ECONOMIST: The peace process is still on track, but it's a slow one

Officials are still trying to determine what caused the fire that lead to the explosion.

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Investigators were trying to determine what caused 114 barges to break loose in St.

NPR: More Rain Expected For Already Swollen Rivers