The evidence comes from a 40, 000-year-old human fossil with delicate toe bones indicative of habitual shoe-wearing, experts say.

专家们说证据来自于一个有4万年历史的人类趾骨化石，这个纤细的趾骨化石是惯常穿鞋的象征。

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In a previous study, Trinkaus found that shoe-wearing and barefoot human groups show characteristic differences in the size and strength of their middle toe bones.

Trinkaus在先前的研究中发现，穿鞋和赤足人群的脚部特征存在差异，具体体现在他们中趾骨的大小和力量上。

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And CAT scans of Tut turned up foot deformities, like a missing toe bone, and bone necrosis, which means some of his foot bones were dying due to poor blood circulation.

而CAT扫描则显示图特王脚部畸形，像是没有趾骨，并且伴有骨坏死，也就是说他的脚部骨头因为血液循环不畅而坏死。

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Numerous other dodo parts were also unearthed, such as skull fragments, beak bones, vertebrae, wing bones and toe bones.

BBC: NEWS | Science/Nature | Dodo skeleton find in Mauritius