He tested his idea in a tiny nematode worm called Caenorhabditis elegans and, in 1994, published a paper showing how touch-receptor proteins are distributed around this worm.

他于1994年在一种称为秀丽隠杆线虫的小型线虫中试验了这一想法，标示出接触受体是如何在整个虫体中分布的，并据此发表了一篇文章。

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He tested his idea in a tiny nematode worm called Caenorhabditis elegans and, in 1994, published a paper showing how touch-receptor proteins are distributed around this worm.

ECONOMIST: The 2008 Nobel science prizes