The former Spanish colony struck oil in the 1990s and is sub-Saharan Africa's fourth-largest exporter after Angola, Nigeria and Sudan.
这个前西班牙殖民地于二十世纪九十年代勘探出石油，现在俨然是撒哈拉以南非洲大地上第四大石油输出国，紧随安哥拉、尼日利亚和苏丹之后。
It makes the whole ant colony have a better balance in the "exploration" and "learning" to adjust the three sub-class sizes and the moving step lengths.
通过自适应地调整三个子类蚁群的规模和搜索步长，使得整个蚁群在“探索”和“学习”之间处于一个较好的平衡。
The sub stitue reproductive colony can produce substitute reproductive once more, and then there may appear substitute reproductive in wing-pad form.
在补充生殖蚁群体中，能再次产生补充生殖蚁，再生补充生殖蚁的品级可出现翅芽型补充生殖蚁。
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