The exospore is smooth and the surface ornamentation is formed by the perispore.
外壁光滑。 周壁较厚，由周壁形成孢子的表面纹饰。
The perispore can be distinguished into three layers, i. e. an outer perispore layer, a middle perispore layer and an inner perispore layer.
周壁是由绒毡层残余物在外壁表面沉积形成， 可分为周壁内层、周壁中层和周壁外层三部分。
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