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McGeechan has chosen a proven combination, all from the Ospreys, in the back three, but the partnerships of Roberts and Earls at centre, O'Gara and Blair at half-back, and Shaw and O'Connell at lock, will be tried for the first time.
BBC: Earls given nod for Lions opener
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Ryan Caldwell replaces Mick O'Driscoll at lock, where he will partner Devin Toner with Dan Tuohy of Ulster named at blindside flanker.
BBC: Mike Blair continues his recovery with Scotland A
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Ireland put in a remarkable 99 tackles, missing just one, and lock O'Connell said he felt both sides played as if they wanted to make amends for their most recent performances.
BBC: Martin Johnson ponders England approach