Nucleolar structure is one of the most obvious symbols. However, little is known about its evolutionary origin. Some previous studies implied that Giardia has no nucleolus.

核仁是真核细胞最显著的标志之一，但关于其进化上的起源迄今仍所知甚少，已发现有迹像表明原始性真核生物贾第虫没有核仁。

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These changes are discussed in relation to current knowledge about nucleolar structure as well as the organization of chromosomes and transcription factories in nuclear regions outside the nucleolus.

讨论了涉及到有关核仁结构的当前知识的这些变化，以及在核仁外部的细胞核区域中染色体和转录工厂的组织。

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In all the nucleolar cycle, nucleolar FCs and DFCs structure changed.

在整个核仁周期中，核仁的FC和DFC结构组分发生变化。

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