In the marker system while we only say YES one time followed by a reward, we can add encouraging verbal praise during the reward and after the reward.

在响片训练体系中，我们说一次“YES”接着就给狗儿一份奖励，我们可以在奖励的时候和奖励之后增加鼓励的口头赞扬。

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The genetic marker system is of significance in anthropology, forensic medicine and medical genetics.

本遗传标记系统在人类学、法医学和医学遗传学的研究中有重要意义。

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The investigation results of familial members showed that this genetic marker system was in line with Mendelian's law.

家系调查表明这一位点的等位基因按孟德尔规律遗传。

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Certainly, Marker's system doesn't guarantee that a fallen angel will pick itself up off the floor.

FORBES: Timing Is Everything

This "biophony, " as Krause calls it, can be studied as a marker of an eco-system's health.

CNN: Meet the man recording the sound of extinction