并且， 认为凋亡细胞的核小体是主要致病物质（major pathogen） (Kalled, et al. 2001) 。
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major pathogen
主要病原体
以上为机器翻译结果，长、整句建议使用 人工翻译 。
UU was the most major pathogen of NGU.
UU是非淋主要的病原体；
youdao
The major pathogen is Gram-negative bacteria.
主要病原菌是革兰阴性杆菌。
Influenza a virus is the major pathogen not only carrying seasonal influenza but also causing flu pandemic in history.
A型流感病毒不但是季节性流感的主要病原，而且历史上几次流感大流行也都由A型流感病毒引起。
In fact, pathogen detection is bound to be another major component of nanotech in the food industry.
FORBES: Safer And Guilt-Free Nano Foods
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