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It also gives advice on how and when doctors should raise the issue of organ donation.

它对医生何时和怎样对病人提出器官捐赠的问题也给予了指导。

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It will never be known how and when this numeration ability developed, but it is certain that numeration was well developed by the time humans had formed even semi-permanent settlements.

我们永远也不会知道这种计算能力是何时、如何发展起来的，但可以肯定的是，在人类形成半永久定居点的时候，这种计数法已经发展得很好了。

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One of the big issues to discuss is how and when to reduce deficits and economic growth measures as conditions improve.

VOA: special.2010.06.25

But how and when did it happen?

但这是什么时候怎样发生的呢

耶鲁公开课 - 死亡课程节选

He had two brothers and two sisters, and it's amazing, when you read his memoirs, how invisible they are.

他有两个哥哥和两个姐姐，令人惊讶的是当你读他的回忆录时，几乎看不到他们。

耶鲁公开课 - 1945年后的美国小说课程节选

Here's a guide to how and when you may be asked to pay more.

FORBES: Your Next Tax Hike

And how and when do they actually engage with your company (and your competitors)?

FORBES: The Key To Building Customer Relationships: Serve Your Own Team First

They can choose how and when they go about accomplishing the tasks by the set date.

FORBES: 3 Questions To Ask Before Green-Lighting A New Startup Hire