It also gives advice on how and when doctors should raise the issue of organ donation.

它对医生何时和怎样对病人提出器官捐赠的问题也给予了指导。

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It will never be known how and when this numeration ability developed, but it is certain that numeration was well developed by the time humans had formed even semi-permanent settlements.

我们永远也不会知道这种计算能力是何时、如何发展起来的，但可以肯定的是，在人类形成半永久定居点的时候，这种计数法已经发展得很好了。

youdao

Parents must let children know how and when they'll be watching them.

父母必须让孩子们知道他们将在什么时候以何种方式监视他们。

youdao

One of the big issues to discuss is how and when to reduce deficits and economic growth measures as conditions improve.

VOA: special.2010.06.25

But how and when did it happen?

但这是什么时候怎样发生的呢

耶鲁公开课 - 死亡课程节选

He had two brothers and two sisters, and it's amazing, when you read his memoirs, how invisible they are.

他有两个哥哥和两个姐姐，令人惊讶的是当你读他的回忆录时，几乎看不到他们。

耶鲁公开课 - 1945年后的美国小说课程节选

Here's a guide to how and when you may be asked to pay more.

FORBES: Your Next Tax Hike

And how and when do they actually engage with your company (and your competitors)?

FORBES: The Key To Building Customer Relationships: Serve Your Own Team First

They can choose how and when they go about accomplishing the tasks by the set date.

FORBES: 3 Questions To Ask Before Green-Lighting A New Startup Hire