You begin to understand what actions were the mainspring of the story.

你开始明白什么是这个故事的主要情节了。

《柯林斯英汉双解大词典》

Do you begin to understand what inflation does to a society?

通货膨胀对一个社会造成什么样的结果，你开始明白了吗？

youdao

Once you begin to understand this and how it manifests itself, the anger dies down.

只要你开始理解这是怎样表现的，愤怒自然消失了。

youdao

It is only then you begin to understand that this is Northern Ireland during the Troubles.

BBC: Paul Graham: Photographs 1981-2006

Only when you listen that carefully and deeply can you begin to understand and then communicate with another person.

FORBES: Thin-Slicing, Malcolm Gladwell, and the Chinese Character for Listening

It is not until you begin to understand the nature of risk that you learn how fantastic it is as an opportunity creator.

FORBES: Their Risks are Our Opportunities