Because complex species interactions may be involved, identifying a keystone species by removing the species and observing changes in the ecosystem is problematic.

由于其中可能涉及复杂的物种相互作用，通过移除一个关键物种并观察生态系统的变化来确定该物种是有问题的。

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The reef, which in actuality is a collection of thousands of distinct coral reefs, has been designated a World Heritage site for its sheer beauty and uniquely complex and delicate ecosystem.

大堡礁，实际上是一个独特的珊瑚礁群，由于其纯粹的美丽和独特内涵以及微妙的生态系统，已被指定为世界遗产地。

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Since the web went mainstream in 1993, our media "ecosystem", if you like, has become immeasurably more complex.

自1993年网络开始变成主流，我们媒体的“生态系统”，无论你喜欢不喜欢，已变得不可估量的更加复杂。

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To achieve this, decision makers need scientifically credible and independent information that takes into account the complex relationships between biodiversity, ecosystem services, and people.

UNESCO: Social and Human Sciences

Decision makers need scientifically credible and independent information that takes into account the complex relationships between biodiversity, ecosystem services, and people in order to adopt and implement adequate policies.

UNESCO: Social and Human Sciences

The agreement calls for the water system to be treated as a single complex ecosystem made up of countless tributaries, underground aquifers and smaller lakes all held in the Great Lakes basin.

NPR: Governors Meet on Access to Great Lakes