The middle layer cell won't degenerate when the tapetum disintegrates prematurely, This proves the hypothesis that the middle layer cells offer nutrients for the tapetum.

绒毡层的提前解体导致不育系花药的中层细胞不退化，为证明中层为绒毡层供给营养的假说提供了证据。

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Mesoderm --- The middle germ layer in the early embryo; the source of cells that go on to make bones, muscles, connective tissue, heart, hematopoietic system, kidney, and other organs.

原肠胚形成过程中，由上胚层细胞增殖产生的一部分细胞在上、下胚层之间形成的第三层细胞。通常形成骨骼、肌肉、结缔组织、心脏、造血系统、肾脏和其它器官。

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