Traditional settlement may be studied as a complex ecological system.

传统聚落可作为复合生态系统来研究。

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Lake is a complicated, dynamic ecological system, the resumption and management of which is a complex systematic project.

湖泊是一个复杂的动态的生态系统，其恢复与管理是一个十分复杂的系统工程。

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This paper summarized the effects of tree-grass complex system on trees and soil and its ecological and economic benefits.

概述了林草复合生态系统对林木和土壤的影响及林草复合的生态效益和经济效益。

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