其他三种动物的区别在于它们食用的低处枝叶和高处茎杆的比例不同：斑马吃茎杆最多，角马吃叶子最多，而转角牛羚则介于两者之间。

The other three species differ in the proportion of lower leaves and higher stems that they eat: zebras eat the most stem matter, wildebeests eat the most leaves, and topi are intermediate.

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因此，当只有供应不足的质量低劣的食物时，角马，转角牛羚和小羚羊享有了优势。

Thus, when there is only a short supply of poor-quality food, the wildebeest, topi, and gazelle enjoy an advantage.

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