胡柚汁营养丰富，但也有柠檬苦素、柚皮苷等苦味物质。
But naringin and limonin, the main bitter substances in Citrus juice are also determined.
同时也用此方法对四种不同的苦味物质成功地进行了识别。
At last, four different bitter substances are identified successful-ly.
但是苦味物质能够阻碍钙流入气道细胞，所以就能缓解拉紧的组织。
But bitter substances block the channels that allow calcium into cells, which relaxes the tightened tissue.
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