但这件标本没有羽轴，带状尾巴上只有羽枝和小羽枝。

But this specimen lacked the rachis; it just had barbs and barbules down its ribbonlike tail.

youdao

该发现表明，羽毛中的羽枝和小羽枝是先于羽轴进化出来的。

The finding suggests that the barbs and barbules evolved before the rachis in feathers.

youdao

沿着羽轴伸展开去的较小的杆状物叫做羽枝，沿着羽枝伸展开去的更小的细丝则叫小羽枝。

Branching from the rachis are smaller shafts called barbs, and then branching from the barbs are even smaller filaments called barbules.

youdao