由于这些指标与一个国家的财富密切相关，他们以人均国民生产总值为控制变量来看它是否能够单独解释择偶策略的差别。

Since there is a strong correlation between these variables and a country's wealth, they controlled for GNP per head by measuring whether it alone accounted for the difference in mating strategy.

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传统的国民经济核算体系及其主要指标GDP具有一定的局限性，无法反映包括人力资源、自然资产等在内的真实国民财富。

The traditional system of national accounting and its core indicator GDP are limited to reflect the real national welfare including human resource, natural asset and so on.

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