The main effect factors of the quantity change of rodent were "nondensity dependent factor" and the density dependent factors had some effect also.

主要影响因素是非密度制约因素，但密度制约因素亦有一定影响。

youdao

There are as well as density-dependent biological factors such as disease, predation, and intraspecific competition .

还有密度制约的生物因素，诸如病害、捕食和种内竞争等。

youdao

There is no need to regard density-dependent factors as having any special importance.

没有必要认为密度制约因素具有任何特殊的重要性。

youdao