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Dr Lyle Armstrong, of the International Centre For Life at the University of Newcastle Upon Tyne, called the studies a "major development".
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Otters have also been recorded in the centre of major towns and cities, including Birmingham, Glasgow, Newcastle-upon-Tyne and Doncaster.
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Lazily stretching out across the northern perimeter of the city walls, the Villa Borghese looks down upon the Piazza del Popolo and the trio of busy thoroughfares that flow into the centre of town.
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