Clearly, this retroviral gene fills an important niche in mammalian biology, but, on the other hand, mammals were able to reproduce perfectly well before they were first infected.

显然，这种逆转录病毒基因填补了哺乳动物生物学的一个小生境，但是从另一方面来说，在第一次感染此类病毒以前，哺乳动物已经能够完美的生殖了。

youdao

All aspects of your life work in tandem. Success in one area will spill over to another, but on the other hand neglect will destroy your success in any of the other areas of your life.

生命中的每一方面都是协调工作的，在一个领域的成功将有利于其他领域，但是对其他领域的忽视，也将带来霉运。

youdao

I've probably had to read more bad science fiction than anyone else in the world; but, on the other hand, I've probably been privileged to read more good science fiction than anyone else in the world.

很有可能，糟糕透顶的科幻小说我比世上其他任何人都读得多，克从另外一方面看，我读过的优秀科幻小说可能也比世上其他人都多，这真是我的荣幸。

youdao

But,on the other hand, Ms.Teltscher says mobile phone usage has reached sixty-eight percent in developing countries.

VOA: special.2010.10.25

I pride myself on the fact that people who have taken this course find it useful in their subsequent lives, but on the other hand, I think that it's really interesting.

我很骄傲的说,选了这门课的人,会在以后的生活中用到这些知识,而另一方面,我觉着这是门有趣的课

耶鲁公开课 - 金融市场课程节选

and I do like having the book in my hand. But other people, they have the 'Kindles'.

我喜欢有书在手。但其他人，他们有“Kindle” 电子书阅读器。

我的爱好是读书 - SpeakingMax英语口语达人

"On the one hand, the Home Office doesn't want them in but on the other hand, the minister for Europe is saying there is freedom of movement, " he said.

BBC: Airport arrivals sign

Now, of course you could say that's true in Iraq as well, but on the other hand, there are questions about whether U.S. personnel have to be in Iraq, what precautions they need to take.

NPR: The Morality of Negotiating with Kidnappers

But in extending its other, diplomatic, hand to America, China, Russia and others, the North is already seeking to play these neighbours-cum-rivals off against each other.

ECONOMIST: The region is about to get harder to manage, not easier