顺向和逆向演替是边缘型旅游地两种截然不同的动态演替模式。
The favorable and the adverse operating patterns are the two different dynamic operating patterns of the peripheral tourism areas.
建立了两种新的演替模式：恢复式和互补式。
Two new successional models were made: Recovery model and Interreplenishment model.
这种自然演替模式随着滩涂（或沙洲）淤积增高和面积的扩涨，即按高程从低到高的顺序进行。
This mode goes from low to high with the heighten of filling up and enlargement of sandbank (or tidal flat). 3.
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