小孢子败育发生在四分体形成以后。
当小孢子从四分体中释放出来，特殊类型的外壁已经形成。
An extine with its specific pattern had already been formed, when microspores were released from tetrads.
此外，观察了前期I的簇毛麦染色体形态，统计了四分体和小孢子中小麦和簇毛麦染色质微核频率。
Otherwise, the chromosome morphology involving Haynaldia chromatin was observed at preleptotene I, and the frequencies of chromatin nucleus at tetrad stage and at microspore stage were investigated.
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