其它35.3%的QTL没有本身的加性效应，却参与了上位性互作，这些位点可能通过诱发和修饰其它位点而起作用。
The other 35.3 per cent QTLs did not have any significant additive effects of their own but were also involved in epistatic interactions.
但是， 今后的方法应当是以多位点模型、环境互作、上位性检验和多个相关性状联合分析为主。
However, the future approaches for GWAS should be based on the multi-locus genetic model, QTN-by-environment interaction, epistatic detection and multivariate analysis.
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