Milton's been noting that some of the pagan deities that the fallen angels eventually became were male and some were female.

堕落天使化身成的异端神灵,有男有女，弥尔顿本无须说明。

耶鲁公开课 - 弥尔顿课程节选

In the Beginning" can modify the verb "taught" as easily as it can the verb "rose." It can either verb assume, just as Milton's angels can "either Sex assume."

最初“还可修饰动词“宣讲“,也可修饰动词“生出“，它可修饰任意一个动词，正如弥尔顿笔下的天使可以“化男作女“

耶鲁公开课 - 弥尔顿课程节选