The discovery indicates that the upper part of the Relu Formation was formed in dry and hot climate in the Eocene.

该植物孢粉的发现肯定了热鲁组上部地层的形成时代为始新世，亦表明其形成于干旱较热的气候环境。

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Since the beginning of the upper Triassic epoch, the Indian plate upthrusted to the north over the Tethys oceanic crust and led to the closure of the Tethys in Eocene epoch.

晚三迭世开始，印度板块在特提斯洋壳上向北仰冲，并导致特提斯在始新世的关闭。

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Since the beginning of the upper Triassic epoch, the Indian plate upthrusted to the north over the Tethys oceanic crust and led to the closure of the Tethys in Eocene epoch.

晚三迭世开始，印度板块在特提斯洋壳上向北仰冲，并导致特提斯在始新世的关闭。

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