But none of the later fossils considered to be on the direct human line were knuckle-walkers.

不过，在后来出土并被认为与人类进化直接有关的化石上，没有出现与关节行走者有关的证据。

youdao

But none of the later fossils considered to be on the direct human line were knuckle-walkers.

不过，在后来出土并被认为与人类进化直接有关的化石上，没有出现与关节行走者有关的证据。

youdao