潘克塞普曾对老鼠进行了相关研究，他发现，当老鼠玩耍的时候，他们经常发出一种唧唧喳喳很尖的声音——根据科学家表示，这是笑的一种原始形态。
Panksepp has studied rats and found that when they play, they often chirp—a primitive form of laughter, according to the scientist.
潘克塞普所做的研究成果也表明，即使老鼠在玩耍和被搔痒时也会笑，只是声调过高人听不到。
Panksepp's own work concludes that even rats produce a version of laughter in response to play and tickling, with chirps too high-pitched for people to hear.
潘克塞普曾对老鼠进行了相关研究，他发现，当老鼠“玩耍”的时候，他们经常发出一种唧唧喳喳很尖的声音，而这则是笑的一种原始形态。
Panksepp has studied rats and found that when they "play, " they often chirp--a primitive form of laughter, according to the scientist.
潘克·塞普在文章中提出的论点是，动物的笑是人类快乐的基础。
And he makes the argument that animal laughter is the basis for human joy.
华盛顿州立大学的雅克·潘克·塞普专门研究动物的笑声反应，但没有直接参与这项新的研究。他称研究报告令人兴奋不已。
Jaak Panksepp of Washington State University, who studies laughter-like responses in animals but didn't participate in the new work, called the paper exciting.
华盛顿州立大学的雅克·潘克·塞普专门研究动物的笑声反应，但没有直接参与这项新的研究。他称研究报告令人兴奋不已。
Jaak Panksepp of Washington State University, who studies laughter-like responses in animals but didn't participate in the new work, called the paper exciting.
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