从孢子形态上看，栗蕨属与凤尾蕨属有很大差异，将其放在一个科中不合适。
The spore morphology of Histiopteris and pteris is very differential, put genus Histiopteris in family Pteridaceae is not suitable, according to the feature of spore morphology.
此外，有冷杉、云杉、松、铁杉和桦等属乔木花粉和杜鹃科、唇形科、莎草科灌木和草本植物花粉及凤尾蕨、水龙骨、卷柏等属孢子。
Besides, pollen of Abies, Picea, Pinus, Tsuga, Betula, Ericaceae, Labiatae and Cyperaceae and spores of Pteris, Polypodium and Selaginella were also observed.
此外，有冷杉、云杉、松、铁杉和桦等属乔木花粉和杜鹃科、唇形科、莎草科灌木和草本植物花粉及凤尾蕨、水龙骨、卷柏等属孢子。
Besides, pollen of Abies, Picea, Pinus, Tsuga, Betula, Ericaceae, Labiatae and Cyperaceae and spores of Pteris, Polypodium and Selaginella were also observed.
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