Anna Karenina will be Knightley's third collaboration with Wright.
BBC: Keira Knightley to play Anna Karenina in new film
Knightley will play Anna, a married woman who has an affair with a younger soldier.
Macfadyen, best known for his role in the BBC spy drama Spooks, will star opposite Keira Knightley, the Screen Daily film industry website said.
BBC: NEWS | Entertainment | Spy drama star is 'new Mr Darcy'
Scottish actor Macfadyen will appear as the arrogant but eligible Mr Darcy with Knightley as his Elizabeth Bennet.
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