He is in a stable condition at Derriford Hospital, Plymouth, after an operation in Tromso, Norway.

BBC: Scott Bennell-Smith

The University of Tromso team who carried out the research say it is the first study to show women who smoke less than men still get more colon cancer.

BBC: Smoking 'poses bigger risk to women'

Trip leaders, 29-year-old Michael "Spike" Reid, from Plymouth, who shot the bear and Andrew Ruck, 27, who is from Brighton but lives in Edinburgh, left hospital on Monday morning, a spokesman for the University Hospital in Tromso said.

BBC: Scott Bennell-Smith