Although not parents, the pups receive nearly all their food from the escorts for up to three months.
Joanne Roberts, from the zoo, said everyone was pleased to see the pups rehabilitated and back in the wild.
BBC: NEWS | UK | Wales | Mid | Seal pups head for new horizons
The pups were born in September, six months after the zoo opened the UK's first underwater viewing zone for the species.
The pups, which are an endangered species, will grow to a length of 6ft (1.8m) and a weight of around 5st 5lb (34kg).
Everyone pays attention to the pups that emerge from their burrows sometime after January, and the pack takes turns at nursery duty to give the mothers time to find some food for themselves.
BBC: Lonely Planet's top five wildlife parks in South Africa
We all have alternate sources of income, usually JOBS, that support the expenses of producing quality, healthy, well socialized animals, expenses which are virtually never covered by the income from the sale of the few pups the breeder and stud owner do not keep.
FORBES: ASJ: What was Maisey��s life like before you got her?
The four pups, which had been fed with herring, were given the names Mango Chutney, Vindaloo, Kashmir and Halleem by staff.
BBC: NEWS | UK | Wales | Mid | Seal pups head for new horizons
He said the two pups had developed a bond and hoped this would help them in the wild.
Mr Anstell said the seal pups would be suckled for about three weeks, during which time they would put on about 2kg (4.5lb) a day.
The increase was due primarily to the fact that grown pups who would normally leave the pack to seek mates remained with their family unit during severe winters.
Because begging has a cost in terms of energy, Dr Bell speculated that the amount of noise pups make should depend on the degree to which they think they will get a return on their work.
That is, until users teach the robo-pups to recognize new names.
In 1999, the Dutch scientists found 12 seal pups with breathing difficulties, stranded on the coast.
Five European grey wolf pups are the first to be born in the Highlands in 12 years.
Two grey seal pups rescued over the winter have been released back into the wild by a Guernsey animal charity.
Two seal pups, rescued from the waters around Guernsey, are ready to be released back into the wild after being cared for by an animal charity.
Finally, we hauled our sleds to the trailhead in the van, pulling a caravan of yelping pups in a trailer behind us.
Matt Rand, director of Global Shark Conservation for the Pew Environment Group said that this was the latest of a number of studies "confirming the importance of preserving the parts of the ocean where sharks breed and raise their pups".
The warden told how anxious owners had been in touch after pups they believed were legal breeds such as Staffordshire bull terriers grew bigger and bigger.
BBC: NEWS | UK | Northern Ireland | Pit bulls handed over to council
But with the balds weakened, the goldens took up residence and began dining on terrestrial prey as is their wont: feral piglets released by historic human ranching operations on the islands and the tiny island foxes, who only have one to four pups a year.
Pups are less likely to make a noise if they think the returns will be low for example, with carers that are stingy or in bad condition and thus have less food to spare.
应用推荐