With the second row locked up by O'Connell and Jones, that just leaves the back row.
Leinster's in-form Sean O'Brien is selected at number eight with open-side David Wallace completing the back row.
Uthappa stormed down the pitch to swat Dewald Nel high into the back row of the mid-wicket stand.
And with it, he sent a message to Wales coach Warren Gatland who is short of options in the back row.
The 27-year-old will start his second Test alongside experienced pair Williams and Jonathan Thomas in the back row.
Mr Jones (see if you can spot him in the back row) is continuing his (interrupted) tour of Asia.
"I thought they were outstanding in the back row - they were a yard ahead of the piece, " Rea said.
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During the game, he lives and dies with every pass as he watches from the back row of the south stand.
His students thought he was acting, being dramatic about an asinine comment just made by a boy in the back row.
Kieran Campbell starts at scrum-half while Michael Swift is recalled to the second row, with Andrew Browne reverting to the back row.
Despite Ireland's Italian struggles, coach Declan Kidney has made only one change with Jamie Heaslip replacing Denis Leamy in the back row.
The back row of David Wallace, Tom Croft and Jamie Heaslip looks quick and powerful and all are playing in their best positions.
In the back row, skipper Jones moves back to number eight with the Scarlets' Dafydd Jones and the Blues' Robin Sowden-Taylor on the flanks.
England make two further changes because of injury, with Louis Deacon coming into the second row and James Haskell picked in the back row.
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Croft, a star of the Lions tour 12 months ago, returns to the back row after missing the Six Nations with a knee injury.
Llanelli began with a bang when Wyatt, who switched to the back row after Scott Quinnell withdrew injured, powered over from the home side's first attack.
Leamy is the only change in the back row where David Wallace is rested with Jamie Healsip remaining at number eight and Stephen Ferris at blind-side.
Broad-chested and bearded, he sat by himself in the back row of the auditorium, wearing a camouflage trucker hat, a brown polo shirt, and cargo pants.
The Briton will share the back row with Toro Rosso's Sebastian Vettel, who did not take part in qualifying after ploughing into the wall during final practice.
"He's been named as a second row, but I see Mike as a back-five forward who gives me options at lock and in the back row, " added Ruddock.
Gatland has chosen Jones to play in an unfamiliar openside role alongside blindside Jonathan Thomas and number 8 Ryan Jones in the back row for the first test.
"It won't work if they try to outmuscle the Springboks, so I would pick pace in the back row and have both Wallace and Williams in there, " Davies explained.
With Geraint Lewis and Hywel Jenkins gone the back row is weakened, while the loss of Andrew Le Chevalier and Johnny Marsters leaves scant cover in the front row.
Moody's elevation to the captaincy for the first time comes a week after he was dropped for the Murrayfield fixture, before a recall to the back row earlier this week.
Talk to the back row, not the front row.
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London Irish open-side flanker Steffon Armitage will also travel to Paris as cover for the back row, with Johnson stressing that it was a precautionary move because of a number of "niggling injuries".
Centre Graeme Morrison was a stand-out in the two summer clashes with the Pumas but Glasgow's strongest area is the back row, where they can call upon John Barclay, Kelly Brown and Johnnie Beattie.
"It's a real continuity of what she has done in the past, even if it is different because times have changed, " Pene says as he watched the second women's show from the back row.
It's a time-sucking hobby that's contributed to the loss of her job (violinist in the back row of an orchestra) and therapist ("Cyd, you can't grow if you're still immersed in an imaginary social environment").
Other than the enforced absence of Vickery, the other two personnel changes from the win over the United States are in the back row, with Worsley and former captain Dallaglio the players to make way.
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