The LCR Honda team rider was over one minute and twenty one seconds behind winner Aprilia rider Alvaro Bautista.
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Landis, a former mountain biker who toiled for three years as a U.S. Postal Service team support rider for Armstrong, had sought to apply the Texan's conservative meticulous strategy for winning.
Last year, former British Superbikes rider Denning led the Crescent Rizla Suzuki team to the category's manufacturer and rider titles.
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The HTC-Highroad rider, a team-mate of Mark Cavendish, proved too strong for the 2008 winner Fabian Cancellara and Philippe Gilbert.
Noemi captained her team and was its lead rider.
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It was long thought that doping was rife in professional cycling (five-time Tour winner Eddy Merckx once said that the Tour was not won on sandwiches and mineral water), but the eventual admission of France's top rider Richard Virenque and team masseur Willy Voet confirmed it.
Wiggins is now 18th overall and will force his Team Sky bosses to make key tactical decisions as Swede Thomas Lovkvist is now their best-placed rider, 1 min 44 sec ahead of the team leader.
The 33-year-old Wiggins wasn't the only high-profile rider to withdraw, with team Garmin-Sharp's Ryder Hesjedal out of the race after succumbing to illness.
It was another disappointing race for the American Yamaha rider Ben Spies, whose team set him a target of winning two races this season.
Dani Pedrosa was third quickest on his Repsol Honda, while the in-form Colin Edwards (Monster Yamaha Tech 3), Pedrosa's Italian team-mate Andrea Dovizioso and Spanish rider Toni Elias completed the top six.
Sophie Wells, a dressage rider on Britain's equestrian team for the Paralympic Games, calls Pistorius a "brilliant" ambassador for disabled athletes.
That was won by Danish rider Martin Pedersen of the CSC team.
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Two years ago, three cyclists including team leader Tyler Hamilton, a celebrated American rider, were kicked out of cycling after failing drug tests.
And with his team-mate closing the gap in the rider standings, Rossi has plenty to think about ahead of the next race at Phillip Island in Australia on 18 October.
Undoubtedly team boss Roger Legeay will keep faith with the rider seen as France's best bet for individual glory.
Two big German TV companies stopped broadcasting the event after a rider from the home T-Mobile team tested positive early in the race.
The team sprint win came despite the withdrawal of star rider Shane Perkins who pulled out in a show of remorse for making a rude gesture at an official.
Great Britain, which took gold in the team show jumping event this week and its first ever dressage team gold Tuesday, could only manage equal fifth for rider Nick Skelton, disappointing the home crowds.
The call-up sees King's Lynn Stars rider Iversen, 28, return to face the team he left four months ago.
Marco Pantani, who previously used banned drugs, was excluded, as was the Saeco team, whose leader is Laurent Dufaux, the former Festina rider.
Bruyneel's departure comes after RadioShack rider Fabian Cancellara claimed he was close to leaving the team following the report.
The Yamaha rider is on the limit all the time and his team-mate Rossi will be happy if the likes of Stoner and Pedrosa can also challenge Lorenzo.
In other words, Contador, certainly the most talented rider, may land in a less-than-ideal team situation next year.
They'll team up again in the Preakness, with Krigger trying to become the first black rider to win since Willie Simms in 1898.
Rossi's team-mate Colin Edwards was quickest in the two practice sessions, with another Yamaha rider, Carlos Checa, also on a Yamaha, second fastest.
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The 1997 Tour winner has reportedly received offers from other teams since the ban was imposed, but the rider's manager, Wolfgang Strohband, said Ullrich planned to stay with Team Coast.
British rider James Toseland equaled his best finish in MotoGP when he claimed sixth for Edwards' team after Italian Marco Melandri crashed late on.
The CSC team leader took the lead four kilometres from the finish of and finished 12 seconds ahead of Spanish rider Iban Mayo with Dutchman Michael Boogerd third.
Also, former dopers such as rider David Millar and Armstrong's former teammate Jonathan Vaughters, who now runs his own professional team, are working to change the culture which led them all to think they had no choice but to cheat to stand a chance.
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