"The seal pup had made its way into the lorry park via a steep, man-made banking, " he said.

BBC: Smartie the seal

Togo, a one-year-old seal pup who was flown Denmark to his new home at St Andrews Aquarium, on Thursday, was in a separate pool to allow him time to adjust to his new surroundings.

BBC: Seal scales fence to reach pup at St Andrews Aquarium

The team checked the responses of seal mothers by recording the number of pup checks made (where the mother raises her head off the ground and moves it in the direction of her young to check their well-being) during a specific time period.

BBC: Seal pup