Jon Clarke's timely return from an injury-hit campaign at hooker freed Michael Monaghan to play at scrum-half, meaning that Vinnie Anderson was on the bench - and big winter signing Richie Myler missed out on the 17 completely.
Saints clearly missed the inspirational play of veteran scrum-half Sean Long, who failed to make his comeback from a broken jaw.
Scotland's head coach Andy Robinson had hoped the scrum-half would be fit to play at least a part of the two-match Test against Argentina.
France-based Perry Freshwater, who can play on either side of the scrum, has been preferred to Yates as the fourth prop.
The Springboks, inspired by some impressive forward play -- particularly in the scrum -- looked to be cruising to a comfortable win as they led 26-7 with just 12 minutes remaining.
Having joined Gloucester in the summer of 2009 and being able to play on both sides of the scrum, Doran-Jones is among those who could take Somerville's place in the team when he leaves for the Melbourne Rebels in May.
Sunday's match is expected to be decided in the scrum, with the respective front threes set to play the leading role.
Wigan scrum-half Leuluai insists he is happy to play out of position if it helps New Zealand pull off a shock, even if the 23-year-old has been a half-back virtually his whole career.
"As far as I'm concerned, he's available to play for Wales whenever he needs to play for Wales, " Kingsley Jones told Scrum V.
The scrum-half, 33, who is set to play his last game at Knowsley Road against Wigan, will finish the town centre book signing three hours before kick-off.
But scrum-half Parra made the most of more sloppy Scotland play to extend the lead with a penalty five minutes into the second half.
Treviso demonstrated from the off that they intended to play with pace and width but from the first scrum Leicester began to butcher the home pack and their forward power laid the ground work for the victory.
"The Stade Aime Giral is one of the best places to play rugby, the fans are amazing, " he told BBC Wales' Scrum V.
"I would like to see Hooky play 10 personally - I'd put Jamie Roberts and Jonathan Davies at centre, " Devereux told Scrum V Radio.
Play continued well over the allotted time, however, and when both teams went down for a Springbok scrum on 85 minutes, an Aussie win looked inevitable.
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