He said his organisation, which was founded in the 1980s, had found that communities which were directly involved in conserving wildlife and were also able to earn a living from it were more keen to protect the animals from poachers.

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Ferguson admitted failing to protect the animals from injury, suffering or disease by failing to get them treatment for liver fluke and failing to monitor their welfare.

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Ferguson also admitted failing to protect the animals from injury, suffering or disease by failing to get them treatment for liver fluke and failing to monitor their welfare.

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That charge alleges that Mr Ferguson failed to protect the animals from injury, suffering or disease by failing to get them treatment for liver fluke and failing to monitor the animals.

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The trickle of hard currency brought in by a few intrepid travellers helped buy food from across the border in Botswana and Zambia, and while some rangers sought work in other countries, a hardy band of guides stayed on to try and protect the animals from poaching.

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They covered the animals with thermal blankets, black bags and their own clothes to try to protect them from the chilling winds.

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After his arrest, Mr Sol Cabrera confessed that the animals had travelled in his luggage, and that he had put them under his clothing "to protect them from X-rays" as he was going through customs.

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