But as the 14 men tired, Wasps cranked up the pressure in the closing minutes and won a penalty try from a driving line-out maul which denied them even a losing bonus point.

BBC: Wasps 26-15 Harlequins

The game's opening try came on 15 minutes, veteran prop Budgen emerging with the ball after Exeter had produced a superb driving maul from a line-out on 18 metres.

BBC: Exeter 24-19 Sale

They seemed to be feeling the effects of the quick turnaround and Tonga made an explosive start with Toulouse's Maka going over after a rolling maul from a line-out after 72 seconds.

BBC: USA 15-25 Tonga

Johnson was a product of the amateur age who adapted seamlessly to professional rugby and was peerless in the tight, excelling in the scrum and rolling maul - not to mention the line-out.

BBC: Who is Courtney Lawes?

They will also clamp down on players who clear out opponents not involved in a ruck or maul.

BBC: Zero tolerance on mauls and rucks

Royal XV skipper Willhelm Koch stretched out for a close-range try before hooker Rayno Barnes rounded off an impressive driving maul to give their side the 15-point margin before O'Gara's inside pass set up wing Tommy Bowe to cross.

CNN: Lions limp to victory in South African opener