She was not identified until DNA matched samples from her family 18 years later.

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Some 7, 591 of these were subsequently matched with samples from crime scenes, including those from 88 murders and 116 rapes.

ECONOMIST: DNA evidence

Lloyd was traced when DNA from his sister, taken when she was arrested on an unrelated matter, was matched to samples taken at the time of the offences.

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However, only one of the three positive samples actually matched those taken from people who had been diagnosed with the clinical disease.

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Reports said police identified Ziya T. as the main suspect in the case after DNA samples taken from his two brothers and a sister matched those found beneath Sierra's nails.

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