For societies that survive on hunting, gathering and fishing, the loss of legal rights over land they may have lived on for centuries is particularly hard.
In 2010, Romney had plenty of passive-loss carryovers, so he is not trying to milk every loss on his return by hunting for passive-activity income either.
Prof Lee added that human-induced pressures, such as hunting, along with projected climate change and habitat loss, have the potential to change the dynamics of elephant populations and, jeopardise the long-term future of the species.