Non Evans goes from full-back to wing and Naomi Thomas fly-half to full-back.
The 24-year-old, who joins on a two-year deal, can play either full-back or wing.
The only uncapped player is Blues full-back or wing Jamie Roberts, who could make his debut against England.
The 17-stone powerhouse, normally a full-back or wing, has been training in the centre position for his region, with the new season two weeks away.
Whether he plays full-back, wing or centre, he reads the game so well that he finds positions where he can use his speed and agility to score tries.
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And Roberts played full-back and wing before finding a home an inside centre where he will partner the great O'Driscoll in the first Test match in Durban on Saturday.
Emerick, 26, who specialises at centre but can cover full-back and wing, scored the late try that sealed the Dragons' 24-15 Heineken Cup play-off defeat by Overmach Parma on 2 June.
The versatile Sale player can cover full-back, wing and outside centre, and if used as a replacement, could provide more of a cutting edge to a back division that has struggled to create chances.
Smith, who is comfortable at full-back, wing or centre, won the Harry Sunderland Trophy in last year's Grand Final for his man of the match display in Leeds' victory over St Helens and went on to play for England at the World Cup.
England took the inevitable step of moving Monye from full-back to the wing for the second half and England made a dynamic - at least compared to the first half - start.
Woodward will be looking particularly closely at full-back Dan Scarbrough, wing three-quarters Mark Cueto and Phil Christophers, along with the back-row forwards.
Brummer made no mistake with his next attempt, as his fellow centre Robert Ebersohn was sent clear after thrilling interplay from full-back Wilton Pietersen and wing Cecil Afrika.
There is no specialist full-back, with Sale wing Mark Cueto, who ended the Six Nations in the number 15 jersey, the likely starter with Josh Lewsey and Jason Robinson as back-up.
Bowe's burst sliced open the Blues defence and his delicate off-load to full-back-turned-wing Byrne was perfectly timed as the Welsh international dived over for his third Ospreys try of the season.
For both Wigan and Hull, Briscoe had featured at full-back, centre and on the wing, and that experience has given a huge amount to his offensive potency from full-back.
They included versatile Wasps back Josh Lewsey, who was originally named in the squad as a full-back but has been playing at wing and centre of late.
He lines up in a reshaped back division that sees James Hook at full-back, Morgan Stoddart on the other wing and fit-again Jamie Roberts forging a midfield partnership with Jonathan Davies.
McFadden will play in a new back three combination with Keith Earls on the left wing and Luke Fitzgerald at full-back for the first time.
England's Lewsey has played at centre, wing and full-back for Wasps and is currently in his testimonial year after 10 professional seasons with the club.
Kirwan has made two changes from the defeat against Fiji with Tongan-born Christian Loamanu switching from wing to full-back instead of the injured Go Aruga.
Robinson will be able to call on Thompson's versatility, as the grandson of the late Bill McLaren can play at full-back, operate on the wing or play in midfield.
Tight-head prop Adam Jones is set for his 50th cap, while Roberts will be winning his fourth cap in a fourth different position, having already played at wing, full-back and inside centre for his country.
Louise Rickard, with 110 caps behind her at wing and full-back, is on the bench as second row cover while uncapped number eight Vici Owens and centre Adi Taviner make their first appearance on the bench.
The versatile Paterson will be at full-back, having played on the wing in his previous eight appearances for Scotland.
The Exiles were seeking a sixth-straight win but were stunned by three Saracens tries in the first 15 minutes from left wing Ratuvou, full-back Castaignede and fly-half Jackson.
He made 27 starts at full-back, 12 on the right wing, 11 on the left, one at inside centre, two at fly-half and two appearances off the bench.
From a free-kick, the Irishman picked out Mark Wilson rushing down the right wing and the full-back sent in a perfect low cross which Maloney touched narrowly wide.
The link-man was aware of Priestland steaming up outside him in support and sent the full-back streaking away down the right wing with a perfectly timed pass, before Jones kept perfection intact with a touchline conversion.
With Ashton viewing him as a wing rather than full-back, Lewsey loses out to Vainikolo, Strettle, Mark Cueto and Paul Sackey in the 32, with Simpson-Daniel and Tom Varndell - both in the Saxons squad - also pushing hard.
Leinster deserved their victory scoring two first-half tries through Irish wing Shane Horgan and full-back Rob Kearney.
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