The big loser overall was Luxembourg's Frank Schleck, who was caught by his CSC teammate Sastre for three minutes, and slipped from second to sixth overall.

CNN: Sastre keeps yellow jersey in Tour de France

But Frank Schleck of Luxembourg, who finished third overall last year, lost nearly half a minute to Wiggins, coming home 135th at 38 seconds behind teammate Cancellara.

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Saxo Bank, including Luxembourg's Andy and Frank Schleck and Swiss time trial world champion Fabian Cancellara, were third, 12 seconds behind the winners and two seconds off the Liquigas outfit of Italy's Vincenzo Nibali.

BBC: Cavendish in red