That model is the best explanation out there for what happens at scales smaller than the atom, but still has a lot of holes in it, and there are other theories out there that go beyond that model.
Her argument that the selfish-gene model is being superseded by other forms of evolutionary explanation relies on an overinterpretation of those alternatives.
We would further posit that the almost universal current acceptance of this model among cyber security professions also provides an intuitive explanation why so many smart, well-funded people, projects and companies are failing to achieve more meaningful results.