1800年代末，在意大利的里雅斯特，一位名叫西格蒙德·弗洛伊德的医科学生受命研究雄性鳗鱼的睾丸。他假设鳗鱼睾丸是体腔内结成的白质环。

In the late 1800s in Trieste, Italy, a medical student named Sigmund Freud was assigned to investigate the testes of the male eel, postulated to be loops of white matter festooning the body cavity.

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