它们体现了艺术的某种色调变化，没有这些差异就谈不上什么艺术色调。
They express a shade of the art which would be lost without them.
经过硒调色图片的颜色和色调变化在不同的相纸上有不同的表现：从冷调褐色、到紫褐色，有着很细微、可辨别差别。
The colour and degree of tonal changes achieved will vary according to the type of paper: ranging from cool chocolate-brown, through purplish brown, to little discernable change.
皮肤被毛必须为一个完整的颜色，但可以有相同颜色的色调变化，可以为黑色、深色或浅灰色，或银白色，浅黄褐色或奶油色。
The coat must be of one over-all color at the skin but may be of varying shades of the same color in the full coat, which may be black, blue, dark or light grey, silver platinum, fawn or cream.
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